Biological Sciences
1 Harvard University
United States
2 Massachusetts Institute of Technology (MIT)
United States
3 University of Cambridge
United Kingdom
4 University of Oxford
United Kingdom
5 Stanford University
United States
6 University of California, Berkeley (UCB)
United States
7 Yale University
United States
8 California Institute of Technology (Caltech)
United States
9 University of California, Los Angeles (UCLA)
United States
10 University of California, San Diego (UCSD)
United States
11 Imperial College London
United Kingdom
12 University of Toronto
Canada
13 Johns Hopkins University
United States
14 Columbia University
United States
15 The University of Tokyo
Japan
16 Cornell University
United States
17 University of Pennsylvania (UPenn)
United States
18 McGill University
Canada
19 Princeton University
United States
20 University of Chicago
United States
21 Arizona State University
United States
22 ETH Zurich (Swiss Federal Institute of Technology)
Switzerland
23 University of Michigan
United States
24 University of British Columbia (UBC)
Canada
25 Duke University
United States
26 University of Melbourne
Australia
27 University of California, San Francisco
United States
28 National University of Singapore (NUS)
Singapore
29 Kyoto University
Japan
30 Rockefeller University
United States
31 University of Washington
United States
32 University of Edinburgh
United Kingdom
33 Tokyo Metropolitan University
Japan
34 UCL (University College London)
United Kingdom
35 Australian National University (ANU)
Australia
36 Pennsylvania State University (Penn State)
United States
37 Washington University in St. Louis
United States
38 University of Queensland (UQ)
Australia
39 University of Auckland
New Zealand
40 University of Sydney
Australia
41 King's College London (KCL)
United Kingdom
42 University of California, Davis (UCD)
United States
43 Peking University
China
44 New York University (NYU)
United States
45 Karolinska Institute
Sweden
46 University of Hong Kong (HKU)
Hong Kong
47 University of Manchester
United Kingdom
48 Ruprecht-Karls-Universität Heidelberg
Germany
49 University of Wisconsin-Madison
United States
50 Osaka University
Japan